Sabo, the Chief of Staff of the Revolutionary Army, contacts Koala and everyone in Kamabakka Queendom. He explains to them about the things that he is accused of, but someone who was eavesdropping on the call takes a stunning action.
Luffy and Chopper fall into the sea along with the mysterious girl. Jimbei tries to help them but is attacked by the unknown creature. Meanwhile, the Sunny is hit by torpedoes and the rest of the Straw Hats are thrown into the sea.
After the stranger identifies herself as Dr. Vegapunk, she's tasked by another individual to bring the crew aboard the Sunny to the lab. Luffy's group explores around Egghead and meets another individual who identifies herself as Vegapunk.
While Luffy and the others are enjoying the futuristic machines of Egghead, someone who resembles Bartholomew Kuma comes after them. When Luffy tries to fight back, Bonney stops him and reveals her shocking relationship with Kuma.
When Blackbeard bumps into Law, strange events begin to happen, a contagious disease, a falling mountain, a battle between pirate crews, including the two captains. Who's Blackbeard's prisoner? Who will win and take the Road Poneglyphs?
Bonney changes the ages of Luffy, Chopper, Jinbe, and hers, so that the Pacifista won't recognize them. Meanwhile, Lilith gives the rest of the crew a tour, and they respond enthusiastically, until someone who looks like Jinbe show up.
Nami and the others are attacked by a Pacifista who resembles Jimbei and while they're in battle, some of Vegapunk's Satellites look on with excitement. Meanwhile, Luffy and the others find something extraordinary at the scrap heap.
Vegapunk Shaka shares with Robin and the others his theory about an ancient kingdom and the 100-Year Void. He even says that he visited Ohara after its destruction. Meanwhile, Luffy and Chopper try to activate the giant robot.
Shaka reveals the secret behind the foundation of the Revolutionary Army and how the relationship between Vegapunk and Dragon affected it. Meanwhile, Vegapunk explains to Luffy and the others about his special power and how his clones work.
Vegapunk reveals the shocking history of the ancient giant robot and asks Luffy to take him off Egghead. Rob Lucci and his fellow Cipher Pol agents arrive at the island and ask for permission to enter the port.
CP-0 makes a forced landing at Egghead and people on the Fabriophase are plunged into confusion while Shaka and the others prepare to fight back on the Labophase. The enraged Atlas stands up to defend the peace of the island.
Luffy transforms into the Warrior of Liberation as Rob Lucci also shows his awakened Devil Fruit form. As the two clash, Vegapunk reveals the shocking truth of the Gum-Gum Fruit and his eye-opening theory on Devil Fruits.
Vegapunk describes how he managed to grant the Seraphim Devil Fruit powers, which surprises Franky and the others. Meanwhile, Luffy is still facing Lucci in his savage, awakened form, but the rocket to the Labophase is about to leave.
Vegapunk asks the Straw Hats to take him on board their ship which makes Franky and Usopp happy but annoys Nami. Lucci orders his troops to block off all Vegapunk's escape routes and looks to invade the heavily-secured Labophase.
Bepo and Chopper recap Trafalgar Law's involvement in the main story, from the Celestial Dragon Hit-and-Run Incident, the Summit War of Marineford, Law meeting Donquixote Rosinante, the Doflamingo's defeat, and the Raid on Onigashima.